Bertie Carvel (season 6, featured season 5) as Tony Blair

Emerald Fennell (season 4, featured season 3) and Olivia Williams (seasons 5-6) - Camilla Parker Bowles

Claire Foy (seasons 1–2, featured season 4, guest season 5), Olivia Colman (seasons 3–4), and Imelda Staunton (season 5-6) - Queen Elizabeth II

Jared Harris - King George VI (season 1, featured season 2)